*1: Trevaylor
Valley, Penzance, 1861, WC
(PNZ) (Paton 1969a: 755).
*2: Dunmere valley,
Bodmin, 1879, RVT (B) (Paton 1969a:
755).
Forms patches (tall lawns), typically pure and
sometimes several metres across, or grows mixed with other
mainly robust bryophytes. Habitat notes from Cornwall are as
follows. It occurs mainly in moist, shaded sites, less often
in the open, most commonly on acidic rocks close to
water-level, but also on firm soil (silty-loam), or locally on
exposed tree-roots or dead wood. Many records are from
vertical, sloping or horizontal surfaces of granitic or slaty
rocks, shallowly submerged to close above water-level at edges
of small to large streams and rivers (0-2 m above water, but
higher up where rivers flood deeply), in places that are
almost unshaded to rather heavily shaded by deciduous
woodlands. Besides occurrences close to flowing water it
occurs in the upper part of the inundation zone beside
Siblyback Lake (Reservoir). In very humid conditions it
occasionally grows away from water, e.g. well above
flood-levels near the R. Tamar inside deciduous woodland on
soil in flushes and on slaty rock and thin overlying soil of a
N.-facing slope. Associates recorded include Kindbergia
praelonga, Pellia
epiphylla,
Plagiomnium undulatum, Racomitrium
aciculare,
Rhizomnium punctatum, Thamnobryum
alopecurum, less often Isothecium holtii, Solenostoma hyalinum.
An atypical record was from deep inside a shady mine adit high
on a sea-cliff.
Rarely c.fr. (one record at Porthmeor, vc1):
capsules immature 5.