*1: Perranporth sand
dunes, 1935, HHK (BM) (Paton 1969a:
724).
*2: On blown sand
amongst rocks on hillside, 45 m alt., SW. slope of Brea Hill,
SW97, DTH 02-001 (BBSUK) (Rothero 2003:
51).
Grows as low lawns, smaller patches or intermixed
with other low mosses. Notes on habitats in Cornwall are as
follows. Most records are from short dune grassland developed
over calcareous sand, on flat ground or slopes among stable
dunes, beside paths, at edges of dune-slacks, or in turf
developed on blown sand on cliff tops or coastal slopes. It is
often locally common in these habitats in vc1 and appears to
have previously been much overlooked because of its rather
nondescript appearance. These sites are fully insolated and
have free-draining substrates that are dry for long periods. A
few records from atypical sites near coasts on thin sandy soil
accumulations over hard substrates: once over old horizontal,
mortared brickwork at edge of dunes and two records from thin
sandy layers over old copper mine-spoil. Other atypical
records include tall dense patches growing with other mosses
on thin soil over old unshaded concrete of disused railway
platform inland (SE. of Connor Downs), small amounts in
crevices of old tarmac of road through fixed dunes (Penhale
Camp), and small amount in very low vegetation in open sandy
patch near low serpentinite outcrop in short coastal heath
(Kynance). Associates include Amblystegium serpens
var. salinum,
Barbula convoluta,
Bryum cf. algovicum, Didymodon fallax, Didymodon insulanus,
Didymodon
tophaceus, Didymodon vinealis, Ditrichum gracile, Pleurochaete
squarrosa, Pseudocrossidium
hornschuchianum, Trichostomum
crispulum, Aphanes sp., Cerastium sp., Plantago
coronopus.
No vegetative propagules seen. Single record c.fr.
(Brea Hill, vc2): capsules immature 1.