*1: Chyenhal Moor,
Newlyn, 1862, WC (PNZ) (Paton 1969a:
716).
*2: Withiel, 1871, RVT
(B) (Paton 1969a: 716).
Habitat notes from Cornwall are as follows.
Colonist of acidic soil in mainly damp places, often on clayey
substrates, but also on gravelly, sandy, humic or peat
surfaces. Commonly unshaded or lightly shaded, but also quite
well shaded at times, e.g. in Grey Willow scrub. Recorded from
variety of habitats with disturbed or recently exposed soil,
including arable fields (stubble), exposed soil on bank in
meadow, cattle-trampled area in marshy pasture, stream and
river banks, clayey bank beside lake (The Loe), ditch-banks
and other banks, tracks in or beside woodland and scrub, soil
heaps beside damp woodland track, wood edges and a woodland
clearing, china-clay spoil, damp clay overlying concrete
steps, and firm soil, peat or sediments exposed by falling
water of reservoirs (where it may be abundant). Associates
recorded include Anthoceros
punctatus,
Ceratodon purpureus, Dicranella
heteromalla,
Trichodon cylindricus, Epipterygium
tozeri,
Fossombronia pusilla, Lunularia cruciata, Nardia scalaris, Pellia epiphylla, Phaeoceros laevis, Pogonatum aloides, Pohlia annotina, Pseudephemerum
nitidum, Tortula
truncata; rarely also Dicranella crispa, Ephemerum serratum, Fossombronia
wondraczekii, Pohlia camptotrachela.
Found with young plants of Crassula helmsii and
Littorella uniflora
at reservoir edges and with Fissidens incurvus on
lake-edge bank.
Rhizoidal tubers seen several times (but not often
searched for). Frequently c.fr.: capsules immature 1-3, 8, 10,
11; dehiscing 2, 11, dehisced 1,
3.