*1: Penzance, 1841, JR
(E) (Paton 1969a: 692).
*2: Marsh E. of Red
Down, W. of Launceston, Aug. 1961, JAP
(BBSUK) (Paton 1962: 357, 1969a:
Under-recorded in past because male plants not
identified (at least up to March 1999) and female plants only
recorded when perianths present.
Habitat notes from C&S are as follows. On peaty
substrate in areas of mires open due to cattle or
horse-grazing and trampling. Unshaded acidic flushes (near Philonotis fontana, Chrysosplenium
of small streams in mires, slightly to moderately shaded by
Grey Willows and in well-grown GreyWillow
carr. Associates include Aneura pinguis, Brachythecium
succulentum. Wet soil at edges of small stream in Grey
Willow carr; soil shaded on
floor of wet Grey Willow carr. On firm soil of
low wet bank in flush above sea-cliff. On water of ruined wall
of china clay dry, trickling with water and shaded by Grey
Willows (near Pellia
endiviifolia). Flushes on slopes of old china clay spoil,
in or beside recently abandoned quarries and spoil-heaps,
unshaded or partly under Juncus effusus [male
plants] (with Aneura
rivulare, Philonotis fontana).
Edge of pool in china-clay spoil. Damp soil of cattle-trodden
marshy pasture near stream, slightly shaded by deciduous
trees. Base of pathside bank near stream, part shaded
deciduous trees. Damp edge of track, part shaded by edge of
Antheridia well developed: 1, 3. Occasionally ?
c.fr.: capsules immature 3, dehiscing